DAND4, or Cerberus 1, DAN family BMP antagonist, is a human gene encoded by CER1. It may also be known as: Cerberus; Cerberus-related protein; DAN domain family member 4. The encoded protein has an amino acid length of 267 and a mass of 30.1 kDa. Other isoforms may exist. DAND4 is a member of the DAN family. Homologs of this gene exist in other organisms, including: Frog.